[1] Russian Acad Sci, Zool Inst, Univ Skaya Nab 1, St Petersburg 199034, Russia
[2] Tomsk State Univ, Lab Mesozo & Cenozo Continental Ecosyst, Prospekt Lenina 36, Tomsk 634050, Russia
[3] Kazan Fed Univ, Inst Geol & Petr Technol, Kremlevskaya Str 4-5, Kazan 420008, Russia
[4] St Petersburg State Univ, Vertebrate Zool Dept, Univ Skaya Nab 7-9, St Petersburg 199034, Russia