[1] JAMSTEC, Dept Biogeocher, Yokosuka, Kanagawa, Japan
[2] Univ Granada, Dept Estratig & Rakontol, Granada, Spain
[3] JAMSTEC, Res & Dev Ctr Global Change, Yokosuka, Kanagawa, Japan
[4] Goethe Univ Frankfurt, Inst Geosci, Frankfurt, Germany
[5] Heidelberg Univ, Inst Earth Sci, Heidelberg, Germany
[6] JAMSTEC, Res & Dev Ctr Submarine Resources, Yokosuka, Kanagawa, Japan
[7] JAMSTEC, Dept Solid Earth Geochem, Yokosuka, Kanagawa, Japan
[8] Fukushima Coll, Natl Inst Technol, Dept Chem & Biochem, Iwaki, Fukushima, Japan