[1] Univ Paris Saclay, Univ Paris Sud, CNRS, Ecol Systemat Evolut,AgroParisTech, F-91400 Orsay, France
[2] UCL, Ctr Biodivers & Environm Res, Dept Genet Evolut & Environm, Darwin Bldg,Gower St, London WC1E 6BT, England
[3] Univ Lausanne UNIL Sorge, Biophore, Dept Ecol & Evolut, CH-1015 Lausanne, Switzerland
[4] Leibniz Inst Freshwater Ecol & Inland Fisheries I, Muggelseedamm 310, D-12587 Berlin, Germany
[5] Free Univ Berlin, Inst Biol, Dept Biol, Chem,Pharm, Konigin Luise Str 1-3, D-14195 Berlin, Germany
[6] Berlin Brandenburg Inst Adv Biodivers Res BBIB, Altensteinstr 34, D-14195 Berlin, Germany
[7] Univ Auckland, Sch Biol Sci, Private Bag 92019, Auckland 1142, New Zealand