[1] Res Inst Nat & Forest, B-9500 Geraardsbergen, Belgium
[2] Bundesamt Umwelt BAFU, Abt Wald, Eidgenoss Dept Umwelt Verkehr Energie & Kommunika, CH-3003 Bern, Switzerland
[3] Nat & Forest Agcy, B-1000 Brussels, Belgium
[4] Univ Ghent, Lab Forestry, B-9090 Melle, Belgium
[5] Katholieke Univ Leuven, Dept Earth & Environm Sci, B-3001 Louvain, Belgium