[1] Univ New South Wales, Sch Biol Earth & Environm Sci, Ctr Marine Sci & Innovat, Evolut & Ecol Res Ctr, Sydney, NSW 2052, Australia
[2] Macquarie Univ, Dept Earth & Environm Sci, Macquarie Pk, NSW, Australia
[3] Flinders Univ S Australia, Ecol & Evolut, Coll Sci & Engn, Bedford Pk, SA, Australia