[1] Hebrew Univ Jerusalem, Inst Earth Sci, IL-91905 Jerusalem, Israel
[2] Univ Haifa, Dept Marine Biol, IL-31999 Haifa, Israel
[3] Univ Leeds, Sch Earth & Environm, Leeds, W Yorkshire, England
[4] Univ Calif Berkeley, Dept Environm Sci Policy & Management, Berkeley, CA 94720 USA