[1] Univ Dundee, Ninewells Hosp & Med Sch, Div Pathol & Neurosci, Dundee DD1 9SY, Scotland
[2] Univ London Kings Coll, Guys Hosp, Div Genet & Dev, London SE1 9RT, England
[3] Univ Dundee, Ninewells Hosp & Med Sch, Dept Pathol, Dundee DD1 9SY, Scotland