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Nanjing Univ, Coll Life Sci, Dept Biol Sci & Technol, State Key Lab Pharmaceut Biotechnol,Plant Cell Ph, Nanjing 210093, Peoples R China Nanjing Univ, Coll Life Sci, Dept Biol Sci & Technol, State Key Lab Pharmaceut Biotechnol,Plant Cell Ph, Nanjing 210093, Peoples R China