[1] INRAE, BIOEPAR, Oniris, Site Chantrerie,CS40706, F-44307 Nantes, France
[2] Unite Epidemiol & Bien Etre Porc, Anses Lab Ploufragan Plouzane, Ploufragan, France
[3] Univ Cote dAzur, ISA, CNRS, INRAE, Biot, France
[4] Univ Cote dAzur, Univ Paris Sorbonne, BIOCORE, Inria,INRAE,CNRS, Biot, France
[5] Univ Paris Saclay, MaIAGE, INRAE, F-78350 Jouy En Josas, France
[6] Natl Vet Inst, Dept Dis Control & Epidemiol, S-75189 Uppsala, Sweden