[1] Palau Int Coral Reef Ctr, Koror, Palau
[2] Univ Hawaii, Hawaii Inst Marine Biol, Fisheries Ecol Res Lab, Kaneohe, HI USA
[3] Natl Geog Soc, Pristine Seas, Washington, DC USA
[4] Univ Massachusetts, Dept Environm Conservat, 160 Holdsworth Way, Amherst, MA 01003 USA
[5] Norwegian Res Ctr NORCE, Lab Freshwater Ecol & Inland Fisheries, Bergen, Norway