[1] Rockefeller Univ, Lab Retrovirol, New York, NY 10021 USA
[2] Rockefeller Univ, Aaron Diamond AIDS Res Ctr, New York, NY 10021 USA
[3] Aaron Diamond AIDS Res Ctr, Howard Hughes Med Inst, New York, NY USA
[4] Univ Birmingham, Ctr Syst Biol, Birmingham, W Midlands, England
[5] Univ Glasgow, MRC Ctr Virus Res, Inst Infect Immun & Inflammat, Coll Med Vet & Life Sci, Glasgow, Lanark, Scotland