[1] Cirad, UMR AGAP, F-34398 Montpellier 5, France
[2] Ctr Biotechnol Sfax, Sfax 3018, Tunisia
[3] Univ Montpellier 2, Int Joint Lab Rice Funct Genom & Plant Biotechnol, Nat Key Lab Plant Cell Biotechnol, IRD,UM2,AGI,VAAS,USTH,Agr Genet Inst, Hanoi, Vietnam
[4] King Saud Univ, Coll Food & Agr Sci, Dept Plant Prod, Riyadh 11451, Saudi Arabia