[1] Rutgers State Univ, Inst Marine & Coastal Sci, New Brunswick, NJ 08903 USA
[2] Kunsan Natl Univ, Coll Ocean Sci & Technol, Dept Oceanog, Kunsan, South Korea
[3] Columbia Univ, Lamont Doherty Geol Observ, Palisades, NY 10964 USA
[4] Univ Florida, Dept Fisheries & Aquat Sci, Gainesville, FL USA
[5] Calif Polytech State Univ San Luis Obispo, Dept Biol Sci, San Luis Obispo, CA 93407 USA