[1] Washington Univ, St Louis Childrens Hosp, Dept Pediat, Sch Med, St Louis, MO 63110 USA
[2] Washington Univ, St Louis Childrens Hosp, Dept Mol Biol & Pharmacol, Sch Med, St Louis, MO 63110 USA
[3] Univ Helsinki, Childrens Hosp, Biomedicum, Program Dev & Reprod Biol, Helsinki, Finland
[4] Univ Tampere, FIN-33101 Tampere, Finland
[5] Univ Tampere, Cent Hosp, FIN-33101 Tampere, Finland