[1] Univ Fed Minas Gerais, Inst Ciencias Biol, Dept Bot, Programa Pos Grad Biol Vegetal, BR-31270901 Belo Horizonte, MG, Brazil
[2] Nucleo Pesquisa & Curadoria Herbario, Inst Bot, Sao Paulo, Brazil
[3] Univ Brasilia, Inst Ciencias Biol, Dept Bot, BR-70910900 Brasilia, DF, Brazil