[1] Oregon State Univ, Dept Fisheries & Wildlife, Hatfield Marine Sci Ctr, Newport, OR 97265 USA
[2] Oregon State Univ, Coll Earth Ocean & Atmospher Sci, Hatfield Marine Sci Ctr, Newport, OR 97365 USA
[3] ARS, USDA, Hatfield Marine Sci Ctr, Newport, OR 97365 USA