[1] Jeju Natl Univ, Dept Marine Life Sci, Jeju, South Korea
[2] Jeju Natl Univ, Fish Vaccine Res Ctr, Jeju, South Korea
[3] Kyungpook Natl Univ, Sch Appl Biosci, Daegu, South Korea
[4] Jeju Natl Univ, Interdisciplinary Grad Program Adv Convergence Te, Jeju, South Korea
[5] Jeju Natl Univ, Dept Anat, Sch Med, Jeju, Self Governing, South Korea