[1] Univ Tras os Montes & Alto Douro, Sch Agr & Vet Sci, Dept Vet Sci, P-5001801 Vila Real, Portugal
[2] Univ Tras os Montes & Alto Douro, Ctr Vet & Anim Sci, P-5001801 Vila Real, Portugal
[3] USDA ARS, Beltsville Agr Res Ctr, Anim Parasit Dis Lab, Beltsville, MD 20705 USA
[4] Univ Porto, Inst Biol Mol & Celular, Parasite Dis Grp, Oporto, Portugal