[1] Univ Copenhagen, Dept Geosci & Nat Resource Management, Oster Voldgade 10, DK-1350 Copenhagen K, Denmark
[2] Multiconsult Norge AS, N-5221 Bergen, Norway
[3] Univ Copenhagen, Dept Biol, Freshwater Biol Sect, Univ Pk 4, DK-2100 Copenhagen O, Denmark
[4] Univ Southern Denmark, Biol Inst, Campusvej 55, DK-5230 Odense, Denmark
[5] Aarhus Univ, Dept Geosci, WATEC, DK-8000 Aarhus C, Denmark