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Sun Yat Sen Univ, State Key Lab Biocontrol, Guangdong Prov Key Lab Improved Variety Reprod Aq, Sch Life Sci, Guangzhou 510275, Peoples R China Sun Yat Sen Univ, State Key Lab Biocontrol, Guangdong Prov Key Lab Improved Variety Reprod Aq, Sch Life Sci, Guangzhou 510275, Peoples R China