[1] Yokohama City Univ, Sch Med, Dept Microbiol, Yokohama, Kanagawa 232, Japan
[2] Ehime Univ, Venture Business Lab, Matsuyama, Ehime, Japan
[3] Ehime Univ, Proteosci Ctr, Matsuyama, Ehime, Japan
[4] Natl Hosp Org Nagoya Med Ctr, Clin Res Ctr, Nagoya, Aichi, Japan
[5] Natl Inst Infect Dis, Pathogen Genom Ctr, Tokyo, Japan
[6] Yokohama City Univ, Int Grad Sch Arts & Sci, Yokohama, Kanagawa 232, Japan
[7] Natl Inst Infect Dis, Infect Dis Surveillance Ctr, Tokyo, Japan
[8] Yokohama City Univ, Sch Med, Dept Mol Biol, Yokohama, Kanagawa 232, Japan
[9] Natl Univ Singapore, Dept Microbiol, Singapore 117548, Singapore