[1] Univ Bern, Inst Phys, CH-3012 Bern, Switzerland
[2] Univ Bern, Oeschger Ctr Climate Change Res, CH-3012 Bern, Switzerland
[3] Max Planck Inst Biogeochem, Dept Biogeochem Syst, D-07745 Jena, Germany
[4] Univ Utrecht, Fac Geosci, Dept Earth Sci, NL-3508 TA Utrecht, Netherlands
[5] PBL Netherlands Environm Assessment Agcy, NL-3720 AH Bilthoven, Netherlands
[6] Chinese Acad Sci, Inst Tibetan Plateau Res, Lab Tibetan Environm Changes & Land Surface Proc, Beijing 100101, Peoples R China
[7] Macquarie Univ, Dept Biol Sci, N Ryde, NSW 2109, Australia
[8] Imperial Coll, Grantham Inst Climate Change, Ascot SL5 7PY, Berks, England
[9] Imperial Coll, Div Ecol & Evolut, Ascot SL5 7PY, Berks, England