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Univ Paul Cezanne, Fac Sci & Tech St Jerome, Inst Mediterraneen Ecol & Paleoecol,IRD, Equipe Ecol Microbienne & Biotechnol,CNRS,UMR 611, F-13397 Marseille 20, France Univ Paul Cezanne, Fac Sci & Tech St Jerome, Inst Mediterraneen Ecol & Paleoecol,IRD, Equipe Ecol Microbienne & Biotechnol,CNRS,UMR 611, F-13397 Marseille 20, France