[1] Univ Bourgogne Franche Comte, CNRS, Ctr Rech Climatol, UMR Biogeosci 6282, 6 Blvd Gabriel, F-21000 Dijon, France
[2] AgroSup, Dept Agron Environm, F-21000 Dijon, France
[3] INRA, UMR Ecol & Ecophysiol Forestieres 1137, F-54280 Champenoux, France
[4] Univ Lorraine, UMR Ecol & Ecophysiol Forestieres 1137, BP 239, F-54506 Vandoeuvre Les Nancy, France
[5] CEFE CNRS Montpellier, CREA Ctr Rech Ecosyst Altitude, 67 Lacets Belvedere, F-74400 Chamonix Mt Blanc, France