[1] MRC Mammalian Genet Unit, Harwell, Oxon, England
[2] Univ Cambridge, Addenbrookes Hosp, Dept Med Genet, Cambridge CB2 2QQ, England
[3] Univ Cambridge, Addenbrookes Hosp, Cambridge Inst Med Res, Cambridge CB2 2QQ, England
[4] CSIC UAM, Ctr Mol Biol Severo Ochoa CBMSO, Madrid, Spain
[5] Inst Salud Carlos III, Networking Res Ctr Neurodegenerat Dis CIBERNED, Madrid, Spain