[1] Kyushu Univ, Lab Marine Environm Sci, Fac Agr, Higashi Ku, Hakozaki 6-10-1, Fukuoka 8128581, Japan
[2] Sam Ratulangi Univ, Fac Fisheries & Marine Sci, Manado 95115, Indonesia
[3] Kanazawa Univ, Inst Nat & Environm Technol, Noto Marine Lab, Noto, Ishikawa 9270553, Japan
[4] Niigata Univ, Sado Marine Biol Stn, Fac Sci, Tassha, Sado 9522135, Japan
[5] Tohoku Univ, Grad Sch Agr Sci, Lab Biol Oceanog, Aoba Ku, Amamiya Machi 1-1, Sendai, Miyagi 9818555, Japan
[6] Fukushima Univ, Inst Environm Radioact, Kanayagawai 1, Fukushima, Fukushima 9601296, Japan
[7] Tokyo Univ Marine Sci & Technol, Field Sci Ctr, Tateyama Stn, Sakata 670, Tateyama, Toyama 2940308, Japan
[8] Tohoku Univ, Res Ctr Marine Biol, Grad Sch Life Sci, Sakamoto 9, Asamushi 0393501, Japan
[9] Hokkaido Univ, Usujiri Fisheries Stn, Field Sci Ctr Northern Biosphere, Usujiri 151, Hakodate, Hokkaido 0411613, Japan
[10] Kyoto Univ, Field Sci Educ & Res Ctr, Sakyou Ku, Kyoto 6068502, Japan
[11] Kyushu Univ, Cent Inst Radioisotope Sci & Safety, 744 Motooka, Fukuoka 8190395, Japan