[1] Univ Med Ctr Utrecht, Rudolf Magnus Inst, Dept Neurosci & Pharmacol, NL-3584 CG Utrecht, Netherlands
[2] Univ Utrecht, Rudolf Magnus Inst, Fac Vet Med, Div Behav Neurosci,Dept Anim Sci & Soc, NL-3508 TC Utrecht, Netherlands
[3] Radboud Univ Nijmegen, Dept Organismal Anim Physiol, Nijmegen, Netherlands