[1] Northwest A&F Univ, Coll Forestry, Key Lab Soil Eros & Dryland Farming Loess Plateau, Yangling 712100, Shaanxi, Peoples R China
[2] Chinese Acad Sci, Chengdu Inst Biol, CAS Key Lab Mt Ecol Restorat & Bioresource Utiliz, Chengdu 610041, Sichuan, Peoples R China
[3] Chinese Acad Sci, Chengdu Inst Biol, Biodivers Conservat Key Lab Sichuan Prov, Chengdu 610041, Sichuan, Peoples R China
[4] Chinese Acad Sci, Chinese Journal Appl & Environm Biol, Chengdu Inst Biol, Chengdu 610041, Sichuan, Peoples R China
[5] Univ Quebec, Dept Biol Sci, Ctr CEE ESCER, Montreal, PQ C3H3P8, Canada