[1] Natl Inst Genet, Dept Dev Genet, Mishima, Shizuoka 4118540, Japan
[2] SOKENDAI, Dept Genet, Mishima, Shizuoka 4118540, Japan
[3] Korea Ocean Res & Dev Inst, So Coastal Environm Res Div, Geoje 656830, South Korea
[4] Natl Inst Genet, Ctr Informat Biol, Mishima, Shizuoka 4118540, Japan