[1] Univ London Imperial Coll Sci Technol & Med, Dept Life Sci, Ascot SL5 7PY, Berks, England
[2] Univ Lancaster, Lancaster Environm Ctr, Lancaster LA1 4YQ, England
[3] Univ Toronto Scarborough, Dept Biol Sci, Toronto, ON, Canada
[4] Univ Toronto, Dept Ecol & Evolutionary Biol, Toronto, ON M5S 1A1, Canada
[5] Lincoln Univ, Bioprotect Res Ctr, Canterbury, New Zealand
[6] Newcastle Univ, Sch Biol, Newcastle Upon Tyne NE1 7RU, Tyne & Wear, England