[1] Johns Hopkins Sch Med, Dept Biomed Engn, Baltimore, MD 21205 USA
[2] Vanderbilt Univ, Dept Psychol, Nashville, TN 37240 USA
[3] Univ Colorado, Dept Integrat Physiol, Boulder, CO 80309 USA
[4] Johns Hopkins Sch Med, Dept Neurosci, Baltimore, MD 21205 USA
[5] Univ Colorado, Dept Mech Engn, Boulder, CO 80309 USA