[1] Univ Alberta, Livestock Gentec, Dept Agr Food & Nutr Sci, Edmonton, AB T6G 2P5, Canada
[2] Agr & Agri Food Canada, Lacombe Res & Dev Ctr, 6000 C&E Trail, Lacombe, AB T4L 1W1, Canada
[3] Alberta Agr & Forestry, Lacombe Res & Dev Ctr, 6000 C&E Trail, Lacombe, AB T4L 1W1, Canada
[4] Univ Manitoba, Dept Anim Sci, Winnipeg, MB R3T 2N2, Canada