[1] Mekelle Univ, Dept Biol, Mekelle, Ethiopia
[2] Leiden Univ, Inst Environm Sci, Leiden, Netherlands
[3] Univ Antwerp, Evolutionary Ecol Grp, B-2020 Antwerp, Belgium
[4] Aarhus Univ, Dept Agroecol, DK-4200 Slagelse, Denmark
[5] Mekelle Univ, Dept Land Resource Management & Environm Protect, Mekelle, Ethiopia
[6] Catholic Univ Louvain, Dept Earth & Environm Sci, B-3001 Heverlee, Belgium