[1] Univ Estadual Campinas, Programa Posgrad Biol Vegetal, Inst Biol, BR-13083862 Campinas, SP, Brazil
[2] Inst Nacl Biodiversidad INBio, Santo Domingo Heredia, Costa Rica
[3] Royal Bot Gardens, Herbarium, Richmond TW9 3AB, Surrey, England
[4] DIPEQ, Inst Pesquisas Jardim Bot Rio de Janeiro, BR-22460030 Rio De Janeiro, RJ, Brazil
[5] Univ Estadual Campinas, Dept Biol Vegetal, Inst Biol, BR-13083862 Campinas, SP, Brazil