Podolasiini Howden, new tribe, and a revision of the included genera, Podolasia Harold and Podostena Howden, new genus (Coleoptera: Scarabaeidae: Melolonthinae)

A new tribe, Podolasiini Howden, is proposed for two closely related melolonthine genera: Podolasia Harold and Podostena Howden, new genus. Major tribal characters are: size small (total length less than 10 mm); mouthparts greatly reduced, hidden under clypeus; clypeus elevated basally; antenna 8- or 9-segmented; antennal club 3-segmented, small, oval; abdominal sternites 2-5 connate medially; tarsal claws simple, slender, posterior claws equal;females flightless. Podolasia Harold includes nine species, of which four are new: P. lavignei Howden, new species, from San Luis Potosi, P. stillwellorum Howden, new species, from Texas, P. longipenis Howden, new species, from Durango, and P. rotundipenis Howden, new species, from Durango and Chihuahua. Podolasia saylori Howden is placed as a junior synonym of P. peninsularis Howden. Podostena Howden, new genus, includes the type species, Pt. bottimeri (Howden), and Pt. rileyi Howden, new species, from Texas, Pt. sleeperi Howden, new species, and Pt. litoralis Howden, new species, both from Baja California Sur. The first known female of a species in either genus, Podostena bottimeri, is described. All species of both genera are keyed and illustrated. Relationships within Podolasia and its out group, Podostena, are discussed and the geographic patterns are mapped.