[1] Natl Univ Singapore, Temasek Life Sci Lab, Singapore 117604, Singapore
[2] Natl Univ Singapore, Dept Biol Sci, Singapore 117543, Singapore
[3] Nara Inst Sci & Technol, Ikoma, Nara 6300192, Japan
[4] Cold Spring Harbor Lab, Cold Spring Harbor, NY 11724 USA
[5] Nagoya Univ, Grad Sch Sci, Div Biol Sci, Chikusa Ku, Nagoya, Aichi 4648602, Japan
[6] Nagoya Univ, JST ERATO Higashiyama Live Holon Project, Chikusa Ku, Nagoya, Aichi 4648602, Japan