[1] Univ St Andrews, Scottish Oceans Inst, East Sands Fife, St Andrews KY16 LB, Fife, Scotland
[2] Univ South Wales, Sch Appl Sci, Cemetery Rd, Glyntaff CF37 4BD, Pontypridd, England
[3] Univ St Andrews, Sch Math & Stat, Ctr Res Environm & Ecol Modelling, The Observ, St Andrews KY16 9LZ, Fife, Scotland
[4] Marine Ecol Consuling, Corkil, Plockton, Scotland
[5] Agroecosyst Lid, Keepers House, Gaick PH21 1NZ, Kingussie Inver, Scotland
[6] Univ Scotland, Carnegie Trust, Andrew Carnegie House,Plitencrieff St, Dunfermline, Fife, Scotland