[1] Univ Helsinki, Bioctr 3, Dept Biosci, FI-00014 Helsinki, Finland
[2] Aalto Univ, Dept Informat & Comp Sci, Adapt Informat Res Ctr, Sch Sci & Technol, Espoo, Finland
[3] Basque Fdn Sci, IKERBASQUE, Bilbao 48011, Spain
[4] Univ Basque Country, Fac Comp Sci, Computat Intelligence Grp, Donostia San Sebastian, Spain
[5] Univ Helsinki, Fac Biol & Environm Sci, Biodivers Conservat Informat Grp, Ctr Excellence Metapopulat Biol,Dept Biosci, FI-00014 Helsinki, Finland