[1] Hiroshima Univ, Grad Sch Biosphere Sci, Higashihiroshima 7398528, Japan
[2] Hiroshima Univ, Grad Sch Integrated Sci Life, Higashihiroshima 7398528, Japan
[3] Univ Western Australia, Sch Biol Sci, 35 Stirling Hwy, Crawley, WA 6009, Australia
[4] Univ Philippines Los Banos, Coll Agr & Food Sci, Inst Crop Sci, Laguna, Philippines
[5] Int Rice Res Inst, DAPO Box 7777, Manila, Philippines
[6] Univ Guelph, Dept Plant Agr, Guelph, ON N1G 2W1, Canada