[1] South China Agr Univ, Coll Hort, Guangzhou 510642, Guangdong, Peoples R China
[2] Guangdong Inst Microbiol, Guangzhou 510070, Guangdong, Peoples R China
[3] Chinese Acad Sci, South China Bot Garden, Key Lab Vegetat Restorat & Management Degraded Ec, Guangzhou 510160, Guangdong, Peoples R China