[1] Univ Birmingham, NIHR Biomed Res Unit, Birmingham, W Midlands, England
[2] Univ Birmingham, Liver Res Ctr, Birmingham, W Midlands, England
[3] Univ Hosp Freiburg, Dept Internal Med 2, Freiburg, Germany
[4] Univ Birmingham, Immun & Infect, Birmingham, W Midlands, England
[5] Univ Jena, Dept Internal Med 4, Jena, Germany
[6] Univ Birmingham, Ctr Human Virol, Birmingham, W Midlands, England