[1] Wageningen Univ & Res, Dept Environm Sci, Wageningen, Netherlands
[2] Univ East Anglia, Sch Environm Sci, Norwich, Norfolk, England
[3] Univ Rennes, Geosci Rennes, Rennes, France
[4] Silixa Ltd, Elstree, Herts, England
[5] Univ Birmingham, Sch Geog Earth & Environm Sci, Birmingham, W Midlands, England
[6] Oregon State Univ, Biol & Ecol Engn, Corvallis, OR 97331 USA