[1] Univ Durham, Dept Earth Sci, Palaeoecosyst Grp, Durham DH1 3LE, England
[2] Lund Univ, Dept Geol, SE-22362 Lund, Sweden
[3] Northwest Univ, State Key Lab Continental Dynam, Shaanxi Key Lab Early Life & Environm, Xian 710069, Shaanxi, Peoples R China
[4] Northwest Univ, Dept Geol, Xian 710069, Shaanxi, Peoples R China
[5] Swedish Museum Nat Hist, POB 50007, SE-10405 Stockholm, Sweden
[6] Univ Lille, CNRS, UMR Evo Eco Paleo 8198, F-59000 Lille, France
[7] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, CAS Key Lab Econ Stratig & Palaeogeog, 39 East Beijing Rd, Nanjing 210008, Jiangsu, Peoples R China
[8] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, Ctr Excellence Life & Palaeoenvironm, 39 East Beijing Rd, Nanjing 210008, Jiangsu, Peoples R China