[1] RDA, Natl Inst Agr Sci, Natl Agrobiodivers Ctr, Jeonju 54874, South Korea
[2] RDA, Int Technol Cooperat Ctr, Jeonju 54875, South Korea
[3] RDA, Dept Southern Area Crop Sci, Upland Crop Breeding Div, Natl Inst Crop Sci, Miryang 50424, South Korea
[4] Jeonbuk Natl Univ, Dept Food Sci & Technol, Jeonju 54896, South Korea
[5] Jeonbuk Agr Res & Extens Serv, Iksan 54591, South Korea