[1] Univ La Reunion, Lab ECOMAR, St Denis 97715 9, Reunion, France
[2] CEBC CNRS, UPR 1934, F-79360 Beauvoir Sur Niort, France
[3] Univ Lisbon, Ctr Estudos Ambiente & Mar CESAM, Museu Nacl His Nat, P-1250102 Lisbon, Portugal
[4] Univ Coimbra, Dept Life Sci, Inst Marine Res IMAR CMA, P-3000 Coimbra, Portugal
[5] Ctr Environm & Educ, Roche Caiman, Mahe, Seychelles