[1] Huazhong Agr Univ, Natl Key Lab Crop Genet Improvement, MOA Key Lab Crop Ecophysiol & Farming Syst Middle, Coll Plant Sci & Technol, Wuhan 430070, Hubei, Peoples R China
[2] Henan Agr Univ, Coll Tobacco Sci, Zhengzhou 450002, Henan, Peoples R China
[3] Yunnan Tobacco Res Inst, Yuxi 653100, Yunnan, Peoples R China