[1] Univ Fed Goias, ICB, Dept Biol Geral, BR-74001970 Goiania, Go, Brazil
[2] Univ Brasilia, Programa Posgrad Biol Anim, BR-70910900 Brasilia, DF, Brazil
[3] Univ Fed Goias, Programa Posgrad Ciencias Ambientais, BR-74001970 Goiania, Go, Brazil
[4] Univ Estadual Maringa, BR-87020900 Maringa, Parana, Brazil
[5] Univ Fed Rio de Janeiro, Rio De Janeiro, Brazil
[6] Univ Connecticut, Storrs, CT USA
[7] Jaguar Conservat Fund, Mineiros, Go, Brazil
[8] Univ Estadual Goias, Unidade Univ Anapolis, Goias, Go, Brazil