[1] Univ Tras Os Montes & Alto Douro, Anim & Vet Res Ctr, Anim Prod Grp, P-5001801 Vila Real, Portugal
[2] INRA, Ctr Res Clermont Ferrand, F-63122 St Genes Champanelle, France
[3] Univ Tras Os Montes & Alto Douro, Ctr Res & Technol Agroenvironm & Biol Sci, P-5001801 Vila Real, Portugal
[4] Wageningen Univ, Dept Anim Sci, Anim Nutr Grp, NL-6709 PG Wageningen, Netherlands
[5] Univ Tecn Lisboa, Fac Med Vet, Interdisciplinary Ctr Res Anim Hlth CIISA, Lisbon, Portugal