[1] CSIRO, Australian Natl Insect Collect, POB 1700, Canberra, ACT 2601, Australia
[2] Albany Museum, Dept Freshwater Invertebrates, Somerset St, Grahamstown, South Africa
[3] Rhodes Univ, Dept Zool & Entomol, Grahamstown, South Africa
[4] South African Inst Aquat Biodivers, ZA-6139 Grahamstown, South Africa
[5] Wild Bird Trust, Natl Geog Okavango Wilderness Project, 20 Loch Ave, ZA-2193 Parktown, South Africa