[1] Cardiff Univ, Sch Earth & Ocean Sci, Cardiff CF10 3AT, Wales
[2] Univ Montpellier, LISAH, INRA, IRD, F-34060 Montpellier, France
[3] Williams Coll, Dept Geosci, Williamstown, MA 01267 USA
[4] Cardiff Univ, Water Res Inst, Sch Biosci, Cardiff CF10 3AT, Wales
[5] Cardiff Univ, Sustainable Places Res Inst, Cardiff CF10 3AT, Wales